Few rifle cartridges have had a greater impact on modern shooting than .223 Remington and 5.56x45mm NATO. These two closely related calibers are the backbone of the AR-15 platform and have become synonymous with semi-automatic rifle ownership in the United States.
Whether used for home defense, target shooting, hunting, or military training, these high-velocity rounds offer precision, versatility, and affordability on a scale unmatched by most other rifle calibers.
The .223 Remington was developed in the late 1950s as a small-caliber, high-velocity cartridge for both civilian and military applications. It gained prominence when the U.S. military adopted a modified version—designated 5.56x45mm NATO—for the M16 rifle during the Vietnam War.
Although the two rounds are dimensionally similar, there are critical differences:
5.56 NATO is loaded to higher pressures
5.56 chambers have a longer throat (leade) for improved cycling under combat conditions
.223 chambers are tighter and may not safely fire 5.56 NATO in all cases
If your rifle is marked “5.56 NATO,” it can safely fire both 5.56 and .223 Remington. However, if it is marked only “.223 Remington,” firing 5.56 NATO can lead to increased chamber pressure and potential reliability or safety issues.
This interchangeability has made the .223/5.56 platform extremely popular among AR-15 owners, offering shooters flexibility across a broad range of loads and purposes.
The AR-15, chambered in .223 Remington or 5.56 NATO, has become the most commonly owned rifle in America. Its success has led to:
Widespread availability of ammo and parts
Huge variety of bullet types and weights
Consistent innovation in barrel, gas system, and optic configurations
These cartridges deliver:
Flat trajectories
Low recoil
Excellent accuracy
…making them ideal for new shooters, youth training, and experienced marksmen alike.
Modern shooters can choose from a wide range of .223 and 5.56 ammo:
55-grain FMJ – Ideal for plinking and general range use
62-grain M855 – Mil-spec “green tip” penetrators for tactical training
69–77 grain match-grade – Designed for long-range precision
Soft point & ballistic tip – Great for varmint and medium game hunting
Bonded hollow points – Optimized for defensive applications
Brands like Federal, Hornady, PMC, Winchester, and Remington offer both bulk options and premium performance loads to suit any shooting purpose.
As long as the AR-15 remains America’s rifle, the .223 Remington and 5.56 NATO cartridges will continue to dominate. Their performance, availability, and adaptability make them the go-to choice for everyone from weekend shooters to tactical professionals.
Whether you're customizing a build, zeroing optics, or stockpiling ammo, these twin calibers offer an unbeatable balance of affordability, reliability, and effectiveness.
Q: Can I shoot 5.56 NATO in a .223 Remington rifle?
A: Not recommended. 5.56 NATO is loaded to higher pressures. It may cause excessive wear or unsafe pressure in rifles marked only for .223 Remington.
Q: What’s the difference between 223 and 556?
A: While they are dimensionally similar, 5.56 NATO is typically loaded to higher pressure, and its chambers are cut differently to handle those pressures safely.
Q: What is the best 5.56 or .223 ammo for self-defense?
A: Look for bonded hollow points, soft point, or ballistic tip rounds such as Hornady TAP, Federal Fusion, or Speer Gold Dot for optimal terminal performance.
Q: Are these rounds good for hunting?
A: Yes, especially for varmints, predators, and small to medium game. Use expanding bullets like soft points or polymer-tipped projectiles for ethical kills.
Q: What grain weight should I use for my AR-15?
A: Most AR-15s are optimized for 55-62 grain bullets, but rifles with faster twist rates (1:7 or 1:8) can handle heavier 69-77 grain match bullets more effectively.